Yes he does. This is the case with all professional photographers unless you sign a contract that copyright is transferred to you.
"The copyright in a photograph belongs to the person who took it - Copyright Designs & Patents Act 1988 - the only exception being employed photographers, where it is his or her employer who owns the copyright unless they have a contractual agreement to the contrary.
In practice, this means that clients may only use photographs taken by a professional photographer in ways that have been agreed at the time they were commissioned. If further uses are required at a later date, permission must be sought from the copyright holder and an additional fee agreed."
http://www.bipp.com/copyright.html