Please Parse

[horseshoes]

1a. Bacteria developing with oxygen with name for tons not doing so.
ANAEROBIC

26a. Including civil engineer, those who conquer pay to use saucy locale.
WORCESTERSHIRE

CE - civil engineer. Hire - pay to use. Worcestershire (sauce). I get this but I'm left with worsters.

Thanks.

[dannyk13]

anagram od bacteria with n replacing t + o(oxygen)

[horseshoes]

Oh for goodness sake!!! I just COULDN'T see it. Thanks Danny.

[donny48]

those who conquer
worst
wəːst/
verb
past tense: worsted; past participle: worsted

get the better of; defeat.
"this was not the time for a deep discussion—she was tired and she would be worsted"
synonyms: defeat, beat, best, get the better of, gain the advantage over, prevail over, triumph over, gain a victory over, trounce, rout, thrash, drub, vanquish, conquer, master, overcome, overwhelm, overpower, overthrow, crush, subdue, subjugate; More

[chiaroscuro]

tr.v. worst·ed, worst·ing, worsts
To gain the advantage over; defeat: was worsted in the argument.

So those who conquer are worsters

[donny48]

Therefore someone who worsted is a worster.

[Togo]

Worst def. transitive verb
:  to get the better of :  defeat
worsters conquer?