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12a From enjoyment of harvest, word for an acorn, berry, drupe, grain, nut, pome or any other edible part of a plant etc (5)
45a Adjective for the sharp biting taste of crab apples, lemons or figurative grapes, or and acidic cocktail of gin or whisky swizzled with citrus juice (4)
56a Ebb and flow of the sea etc (4)
31d ....a word for a wife first, later a female sovereign ruler etc (5)
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